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ccupying 1 370 million cubic kilometres 330 million cubic miles and is the habitat for 11 of known fish species the oceans have a mean depth of 4 000 metres 2 5 miles about 98 of the total water volume is below 100 metres 330 ft and 75 is below 1 000 metres 3 300 ft 2 marine pelagic fish can be divided into coastal inshore fish and oceanic offshore fish coastal pelagic fish inhabit the relatively shallow and sunlit waters above the continental shelf while oceanic pelagic fish inhabit the vast and deep waters beyond the continental shelf even though they also may swim inshore 3 4 pelagic fish range in size from small coastal forage fish such as herrings and sardines to large apex predator oceanic fishes such as bluefin tuna and oceanic sharks 1 they are usually agile swimmers with streamlined bodies capable of sustained cruising on long distance migrations many pelagic fish swim in schools weighing hundreds of tonnes others such as the large ocean sunfish are solitary 1 there are also freshwater pelagic fish in some of the larger lakes such as the lake tanganyika sardine 5 epipelagic fish edit large epipelagic predator fish such as this atlantic bluefin tuna have a deeply forked tail and a smooth body shaped like a spindle tapered at both ends and countershaded with silvery colours small epipelagic forage fish such as this atlantic herring share similar body features as the predator fish described above epipelagic fish inhabit the epipelagic zone the uppermost layer of the water column ranging from sea level down to 200 m 660 ft it is also referred to as the surface waters or the sunlit zone and includes the photic zone the photic zone is defined as the surface waters down to the depth where the sunlight is attenuated to 1 of the surface value this depth depends on how turbid the water is but can extend to 200 m 660 ft in clear water coinciding with the epipelagic zone the photic zone allows sufficient light for phytoplankton to photosynthesize 6 a vast habitat for most pelagic fish the epipelagic zone is well lit so visual predators can use their eyesight is usually well mixed and oxygenated from wave action and can be a good habitat for algae to grow however it is an almost featureless habitat this lack of habitat variation results in a lack of species diversity so the zone supports less than 2 of the world s known fish species much of the zone lacks nutrients for supporting fish so epipelagic fish tend to be found in coastal water above the continental shelves where land runoff can provide nutrients or in those parts of the ocean where upwelling moves nutrients into the area 6 epipelagic fish can be divided broadly into small forage fish and larger predator fish that feed on them forage fish school and filter feed on plankton most epipelagic fish have streamlined bodies capable of sustained cruising on migrations in general predatory and forage fish share the same morphological features predator fish are usually fusiform with large mouths smooth bodies and deeply forked tails many use vision to prey on zooplankton or smaller fish while others filter feed on plankton herring reflectors are nearly vertical for camouflage from the side most epipelagic predator fish and their smaller prey fish are countershaded with silvery colours that reduce visibility by scattering incoming light 6 the silvering is achieved with reflective fish scales that function as small mirrors this may give an effect of transparency at medium depths at sea light comes from above so a mirror that is oriented vertically makes animals such as fish invisible from the side 7 in the shallower epipelagic waters the mirrors must reflect a mixture of wavelengths and the fish accordingly has crystal stacks with a range of different spacings a further complication for fish with bodies that are rounded in cross section is that the mirrors would be ineffective if laid flat on the skin as they would fail to reflect horizontally the overall mirror effect is achieved with many small reflectors all oriented vertically 7 although the number of species is limited epipelagic fishes are abundant what they lack in diversity they make up for in numbers forage fish occur in huge numbers and large fish that prey on them often are sought after as premier food fish as a group epipelagic fishes form the most valuable fisheries in the world 6 many forage fish are facultative predators that can pick individual copepods or fish larvae out of the water column and then change to filter feeding on phytoplankton when that gives better results energetically filter feeding fish usually use long fine gill rakers to strain small organisms from the water column some of the largest epipelagic fishes such as the basking shark and whale shark are filter feeders and so are some of the smallest such as adult sprats and anchovies 8 ocean waters that are exceptionally clear contain little food areas of high productivity tend to be somewhat turbid from plankton blooms these attract the filter feeding plankton eaters which in turn attract the higher predators tuna fishing tends to be optimum when water turbidity measured by the maximum depth a secchi disc can be seen during a sunny day is 15 to 35 metres 9 floating objects edit drifting sargassum seaweed provides food and shelter for small epipelagic fish the small round spheres are floats filled with carbon dioxide which provide buoyancy to the algae great barracuda accompanied by a school of jacks lines of sargassum can stretch for miles along the ocean surface the camouflaged sargassum fish has evolved to live among drifting sargassum seaweed epipelagic fish are fascinated by floating objects they aggregate in considerable numbers around objects such as drifting flotsam rafts jellyfish and floating seaweed the objects appear to provide a visual stimulus in an optical void 10 floating objects may offer refuge for juvenile fish from predators an abundance of drifting seaweed or jellyfish can result in significant increases in the survival rates of some juvenile species 11 many coastal juveniles use seaweed for the shelter and the food that is available from invertebrates and other fish associated with it drifting seaweed particularly the pelagic sargassum provide a niche habitat with its own shelter and food and even supports its own unique fauna such as the sargassum fish 8 one study off florida found 54 species from 23 families living in flotsam from sargassum mats 12 jellyfish also are used by juvenile fish for shelter and food even though jellyfish can prey on small fish 13 mobile oceanic species such as tuna can be captured by travelling long distances in large fishing vessels a simpler alternative is to leverage off the fascination fish have with floating objects when fishermen use such objects they are called fish aggregating devices fads fads are anchored rafts or objects of any type floating on the surface or just below it fishermen in the pacific and indian oceans set up floating fads assembled from all sorts of debris around tropical islands and then use purse seines to capture the fish attracted to them 14 a study using sonar in french polynesia found large shoals of juvenile bigeye tuna and yellowfin tuna aggregated closest to the devices 10 to 50 m farther out 50 to 150 m was a less dense group of larger yellowfin and albacore tuna yet farther out to 500 m was a dispersed group of various large adult tuna the distribution and density of these groups was variable and overlapped the fads also were used by other fish and the aggregations dispersed when it was dark 15 larger fish even predator fish such as the great barracuda often attract a retinue of small fish that accompany them in a strategically safe way skindivers who remain for long periods in the water also often attract a retinue of fish with smaller fishes coming in close and larger fishes observing from a greater distance marine turtles functioning as a mobile shelter for small fish can be impaled accidentally by a swordfish trying to catch the fish 16 coastal fish edit main article coastal fish schooling threadfin a coastal species coastal fish also called neritic or inshore fish inhabit the waters near the coast and above the continental shelf since the continental shelf is usually less than 200 metres deep it follows that coastal fish that are not demersal fish are usually epipelagic fish inhabiting the sunlit epipelagic zone 2 coastal epipelagic fish are among the most abundant in the world they include forage fish as well as the predator fish that feed on them forage fish thrive in those inshore waters where high productivity results from the upwelling and shoreline run off of nutrients some are partial residents that spawn in streams estuaries and bays but most complete their life cycle in the zone 8 oceanic fish edit oceanic fish inhabit the oceanic zone which is the deep open water which lies beyond the continental shelves oceanic fish also called open ocean or offshore fish live in the waters that are not above the continental shelf oceanic fish can be contrasted with coastal fish who do live above the continental shelf however the two types are not mutually exclusive since there are no firm boundaries between coastal and ocean regions and many epipelagic fish move between coastal and oceanic waters particularly in different stages in their life cycle 8 oceanic epipelagic fish can be true residents partial residents or accidental residents true residents live their entire life in the open ocean only a few species are true residents such as tuna billfish flying fish sauries pilotfish remoras dolphinfish ocean sharks and ocean sunfish most of these species migrate back and forth across open oceans rarely venturing over continental shelves some true residents associate with drifting jellyfish or seaweeds 8 partial residents occur in three groups species that live in the zone only when they are juveniles drifting with jellyfish and seaweeds species that live in the zone only when they are adults salmon flying fish dolphin and whale sharks and deep water species that make nightly migrations up into the surface waters such as the lanternfish 8 accidental residents occur occasionally when adults and juveniles of species from other environments are carried accidentally into the zone by currents 8 the huge ocean sunfish a true resident of the ocean epipelagic zone sometimes drifts with the current eating jellyfish the giant whale shark another resident of the ocean epipelagic zone filter feeds on plankton and periodically dives deep into the mesopelagic zone lanternfish are partial residents of the ocean epipelagic zone during the day they hide in deep waters but at night they migrate up to surface waters to feed deep water fish edit scale diagram of the layers of the pelagic zone see also deep sea fish in the deep ocean the waters extend far below the epipelagic zone and support very different types of pelagic fishes adapted to living in these deeper zones 2 in deep water marine snow is a continuous shower of mostly organic detritus falling from the upper layers of the water column its origin lies in activities within the productive photic zone marine snow includes dead or dying plankton protists diatoms fecal matter sand soot and other inorganic dust the snowflakes grow over time and may reach several centimetres in diameter travelling for weeks before reaching the ocean floor however most organic components of marine snow are consumed by microbes zooplankton and other filter feeding animals within the first 1 000 metres of their journey that is within the epipelagic zone in this way marine snow can be considered the foundation of deep sea mesopelagic and benthic ecosystems as sunlight cannot reach them deep sea organisms rely heavily on marine snow as an energy source some deep sea pelagic groups such as the lanternfish ridgehead marine hatchetfish and lightfish families are sometimes termed pseudoceanic because rather than having an even distribution in open water they occur in significantly higher abundances around structural oases notably seamounts and over continental slopes the phenomenon is explained by the likewise abundance of prey species that also are attracted to the structures the fish in the different pelagic and deep water benthic zones are physically structured and behave in ways that differ markedly from each other groups of coexisting species within each zone all seem to operate in similar ways such as the small mesopelagic vertically migrating plankton feeders the bathypelagic anglerfishes and the deep water benthic rattails 17 ray finned species with spiny fins are rare among deep sea fishes which suggests that deep sea fish are ancient and so well adapted to their environment that invasions by more modern fishes have been unsuccessful 18 the few ray fins that do exist are mainly in the beryciformes and lampriformes which also are ancient forms most deep sea pelagic fishes belong to their own orders suggesting a long evolution in deep sea environments in contrast deep water benthic species are in orders that include many related shallow water fishes 19 many species move daily between zones in vertical migrations in the following table they are listed in the middle or deeper zone where they regularly are found species by pelagic zone zone species and species groups include epipelagic 6 mackerel requiem and whale sharks clupeiforms herring anchovy salmonidae salmon atheriniforms flyingfishes halfbeaks sauries perciforms jacks dolphinfish pomfrets barracudas tunas billfish mesopelagic lanternfish opah longnose lancetfish barreleye ridgehead sabretooth stoplight loosejaw marine hatchetfish 20 bathypelagic principally bristlemouth and anglerfish also fangtooth viperfish black swallower telescopefish hammerjaw daggertooth barracudina black scabbardfish bobtail snipe eel unicorn crestfish gulper eel flabby whalefish benthopelagic 6 rattail and brotula are particularly abundant benthic flatfish hagfish eelpout greeneye eel stingray lumpfish and batfish 6 comparative structure of pelagic fishes epipelagic mesopelagic bathypelagic deep sea benthic muscles muscular bodies poorly developed flabby skeleton strong ossified bones weak minimal ossification scales yes none nervous systems well developed lateral line and olfactory only eyes large and sensitive small and may not function variable well developed to absent photophores absent common common usually absent gills well developed kidneys large small heart large small swimbladder vertically migratory fish have swimbladders reduced or absent variable well developed to absent size usually under 25 cm variable species greater than one metre are not uncommon mesopelagic fish edit most mesopelagic fishes are small filter feeders that ascend at night to feed in the nutrient rich waters of the epipelagic zone during the day they return to the dark cold oxygen deficient waters of the mesopelagic where they are ...
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