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ceratophyllaceae wikipedia jump to content main menu main menu move to sidebar hide navigation main page contents current events random article about wikipedia contact us contribute help learn to edit community portal recent changes upload file special pages search search appearance donate create account log in personal tools donate create account log in contents move to sidebar hide top 1 taxonomy toggle taxonomy subsection 1 1 genera 2 references toggle the table of contents ceratophyllaceae 20 languages العربية asturianu corsu čeština dansk esperanto español eesti فارسی français עברית bahasa indonesia 日本語 ქართული latviešu македонски nederlands polski português українська edit links article talk english read edit view history tools tools move to sidebar hide actions read edit view history general what links here related changes upload file permanent link page information cite this page get shortened url switch to legacy parser print export download as pdf printable version in other projects wikimedia commons wikispecies wikidata item appearance move to sidebar hide from wikipedia the free encyclopedia family of aquatic plants ceratophyllaceae temporal range albian recent preꞓ ꞓ o s d c p t j k pg n ceratophyllum submersum scientific classification kingdom plantae clade tracheophytes clade angiosperms order ceratophyllales link 1 family ceratophyllaceae gray genera ceratophyllum ceratostratiotes donlesia ceratophyllaceae is a cosmopolitan family of flowering plants including one living genus 2 commonly found in ponds marshes and quiet streams in tropical and in temperate regions it is the only extant family in the order ceratophyllales 3 species are commonly called coontails or hornworts although hornwort is also used for unrelated plants of the division anthocerotophyta living ceratophyllum grows completely submerged usually though not always floating on the surface and does not tolerate drought taxonomy edit ceratophyllaceae was considered a relative of nymphaeaceae and included in nymphaeales in the cronquist system but research has shown that it is not closely related to nymphaeaceae or any other extant plant family some early molecular phylogenies suggested it was the sister group to all other angiosperms but more recent research suggests that it is the sister group to the eudicots the apg iii system placed the family in its own order the ceratophyllales 1 4 5 the apg iv system accepts the phylogeny shown below 3 angiosperms amborellales nymphaeales austrobaileyales magnoliids chloranthales monocots ceratophyllales eudicots the extinct family montsechiaceae containing the genus montsechia has also been placed in the order ceratophyllales 6 genera edit the family contains one living genus and several extinct genera described from the fossil record including one of the earliest fruit bearing in the form of an achene plants the dakota formation freshwater genus donlesia from early cretaceous 7 ceratophyllum ceratostratiotes donlesia references edit 1 2 angiosperm phylogeny group 2009 an update of the angiosperm phylogeny group classification for the orders and families of flowering plants apg iii botanical journal of the linnean society 161 2 105 121 doi 10 1111 j 1095 8339 2009 00996 x hdl 10654 18083 christenhusz m j m byng j w 2016 the number of known plants species in the world and its annual increase phytotaxa 261 3 201 217 doi 10 11646 phytotaxa 261 3 1 1 2 angiosperm phylogeny group 2016 an update of the angiosperm phylogeny group classification for the orders and families of flowering plants apg iv botanical journal of the linnean society 181 1 1 20 doi 10 1111 boj 12385 kew world checklist of selected plant families permanent dead link angiosperm phylogeny web ceratophyllales gomez bernard daviero gomez véronique coiffard clément barral abel martín closas carles dilcher david l 2020 montsechia vidalii from the barremian of spain the earliest known submerged aquatic angiosperm and its systematic relationship to ceratophyllum taxon 69 6 1273 1292 doi 10 1002 tax 12409 dilcher david l wang hongshan december 2009 an early cretaceous fruit with affinities to ceratophyllaceae american journal of botany 96 12 2256 2269 doi 10 3732 ajb 0900049 pmid 21622341 v t e orders of flowering plants apg iv supergroup plant bryophyta marchantiophyta polypodiophyta acrogymnospermae angiospermae angiosperms amborellales ¹ nymphaeales ¹ austrobaileyales ¹ mesangiosperms chloranthales ceratophyllales magnoliids canellales piperales laurales magnoliales monocots acorales ² alismatales ² petrosaviales ³ dioscoreales ³ pandanales ³ liliales ³ asparagales ³ commelinids arecales poales commelinales zingiberales eudicots ranunculales proteales trochodendrales buxales gunnerales pentapetalae dilleniales superrosids saxifragales rosids vitales fabids zygophyllales celastrales malpighiales oxalidales fabales rosales cucurbitales fagales malvids geraniales myrtales crossosomatales picramniales sapindales huerteales brassicales malvales superasterids berberidopsidales santalales caryophyllales asterids cornales ericales campanulids aquifoliales asterales escalloniales bruniales apiales dipsacales paracryphiales lamiids icacinales metteniusales garryales boraginales gentianales lamiales solanales vahliales ¹ basal angiosperms ² alismatids ³ lilioids category v t e families of flowering plants apg iv supergroup plant rhodophyta chlorophyta marchantiophyta bryophyta anthocerotophyta polypodiophyta acrogymnospermae angiospermae basal angio sperms amborellales amborellaceae nymphaeales hydatellaceae cabombaceae nymphaeaceae austrobaileyales austrobaileyaceae trimeniaceae schisandraceae mesangiospermae magnoliidae canellales canellaceae winteraceae piperales aristolochiaceae saururaceae piperaceae magnoliales myristicaceae magnoliaceae degeneriaceae himantandraceae eupomatiaceae annonaceae laurales calycanthaceae siparunaceae gomortegaceae atherospermataceae hernandiaceae monimiaceae lauraceae chloranthidae chloranthales chloranthaceae lilidae monocots acorales acoraceae alismatales araceae tofieldiaceae alismataceae butomaceae hydrocharitaceae scheuchzeriaceae aponogetonaceae juncaginaceae maundiaceae zosteraceae potamogetonaceae posidoniaceae ruppiaceae cymodoceaceae petrosaviales petrosaviaceae dioscoreales nartheciaceae burmanniaceae dioscoreaceae pandanales triuridaceae velloziaceae stemonaceae cyclanthaceae pandanaceae liliales campynemataceae corsiaceae melanthiaceae petermanniaceae alstroemeriaceae colchicaceae philesiaceae ripogonaceae smilacaceae liliaceae asparagales orchidaceae boryaceae blandfordiaceae asteliaceae lanariaceae hypoxidaceae doryanthaceae ixioliriaceae tecophilaeaceae iridaceae xeronemataceae asphodelaceae amaryllidaceae asparagaceae arecales dasypogonaceae arecaceae commelinales hanguanaceae commelinaceae philydraceae pontederiaceae haemodoraceae zingiberales strelitziaceae lowiaceae heliconiaceae musaceae cannaceae marantaceae costaceae zingiberaceae poales typhaceae bromeliaceae rapateaceae xyridaceae eriocaulaceae mayacaceae thurniaceae juncaceae cyperaceae restionaceae flagellariaceae joinvilleaceae ecdeiocoleaceae poaceae ceratophyllidae ceratophyllales ceratophyllaceae eudicots buxales buxaceae proteales sabiaceae nelumbonaceae platanaceae proteaceae ranunculales eupteleaceae papaveraceae circaeasteraceae lardizabalaceae menispermaceae berberidaceae ranunculaceae trochodendrales trochodendraceae dilleniales dilleniaceae gunnerales myrothamnaceae gunneraceae superrosids saxifragales peridiscaceae paeoniaceae altingiaceae hamamelidaceae cercidiphyllaceae daphniphyllaceae iteaceae grossulariaceae saxifragaceae crassulaceae aphanopetalaceae tetracarpaeaceae penthoraceae haloragaceae cynomoriaceae rosids vitales vitaceae fabids cucurbitales apodanthaceae anisophylleaceae corynocarpaceae coriariaceae cucurbitaceae tetramelaceae datiscaceae begoniaceae fabales quillajaceae fabaceae surianaceae polygalaceae fagales nothofagaceae fagaceae myricaceae juglandaceae casuarinaceae ticodendraceae betulaceae rosales rosaceae barbeyaceae dirachmaceae elaeagnaceae rhamnaceae ulmaceae cannabaceae moraceae urticaceae zygophyllales krameriaceae zygophyllaceae celastrales lepidobotryaceae celastraceae malpighiales pandaceae irvingiaceae ctenolophonaceae rhizophoraceae erythroxylaceae ochnaceae bonnetiaceae clusiaceae calophyllaceae podostemaceae hypericaceae caryocaraceae lophopyxidaceae putranjivaceae centroplacaceae elatinaceae malpighiaceae balanopaceae trigoniaceae dichapetalaceae euphroniaceae chrysobalanaceae humiriaceae achariaceae violaceae goupiaceae passifloraceae lacistemataceae salicaceae peraceae rafflesiaceae euphorbiaceae linaceae ixonanthaceae picrodendraceae phyllanthaceae oxalidales huaceae connaraceae oxalidaceae cunoniaceae elaeocarpaceae cephalotaceae brunelliaceae malvids brassicales akaniaceae tropaeolaceae moringaceae caricaceae limnanthaceae setchellanthaceae koeberliniaceae bataceae salvadoraceae emblingiaceae tovariaceae pentadiplandraceae gyrostemonaceae resedaceae capparaceae cleomaceae brassicaceae crossosomatales aphloiaceae geissolomataceae strasburgeriaceae staphyleaceae guamatelaceae stachyuraceae crossosomataceae geraniales geraniaceae francoaceae huerteales gerrardinaceae petenaeaceae tapisciaceae dipentodontaceae malvales cytinaceae muntingiaceae neuradaceae malvaceae sphaerosepalaceae thymelaeaceae bixaceae cistaceae sarcolaenaceae dipterocarpaceae myrtales combretaceae lythraceae onagraceae vochysiaceae myrtaceae melastomataceae crypteroniaceae alzateaceae penaeaceae picramniales picramniaceae sapindales biebersteiniaceae nitrariaceae kirkiaceae burseraceae anacardiaceae sapindaceae rutaceae simaroubaceae meliaceae superasterids berberidopsidales aextoxicaceae berberidopsidaceae caryophyllales frankeniaceae tamaricaceae plumbaginaceae polygonaceae droseraceae nepenthaceae drosophyllaceae dioncophyllaceae ancistrocladaceae rhabdodendraceae simmondsiaceae physenaceae asteropeiaceae macarthuriaceae microteaceae caryophyllaceae achatocarpaceae amaranthaceae stegnospermataceae limeaceae lophiocarpaceae kewaceae barbeuiaceae gisekiaceae aizoaceae phytolaccaceae petiveriaceae sarcobataceae nyctaginaceae molluginaceae montiaceae didiereaceae basellaceae halophytaceae talinaceae portulacaceae anacampserotaceae cactaceae santalales olacaceae opiliaceae balanophoraceae santalaceae misodendraceae schoepfiaceae loranthaceae asterids cornales nyssaceae hydrostachyaceae hydrangeaceae loasaceae curtisiaceae grubbiaceae cornaceae ericales balsaminaceae marcgraviaceae tetrameristaceae fouquieriaceae polemoniaceae lecythidaceae sladeniaceae pentaphylacaceae sapotaceae ebenaceae primulaceae theaceae symplocaceae diapensiaceae styracaceae sarraceniaceae roridulaceae actinidiaceae clethraceae cyrillaceae ericaceae mitrastemonaceae lamiids icacinales oncothecaceae icacinaceae metteniusales metteniusaceae garryales eucommiaceae garryaceae gentianales rubiaceae gentianaceae loganiaceae gelsemiaceae apocynaceae boraginales boraginaceae vahliales vahliaceae solanales convolvulaceae solanaceae montiniaceae sphenocleaceae hydroleaceae lamiales plocospermataceae carlemanniaceae oleaceae tetrachondraceae calceolariaceae gesneriaceae plantaginaceae scrophulariaceae stilbaceae linderniaceae byblidaceae martyniaceae pedaliaceae acanthaceae bignoniaceae lentibulariaceae schlegeliaceae thomandersiaceae verbenaceae lamiaceae mazaceae phrymaceae paulowniaceae orobanchaceae campanulids apiales pennantiaceae torricelliaceae griseliniaceae pittosporaceae araliaceae myodocarpaceae apiaceae aquifoliales stemonuraceae cardiopteridaceae phyllonomaceae helwingiaceae aquifoliaceae asterales rousseaceae campanulaceae pentaphragmataceae stylidiaceae alseuosmiaceae phellinaceae argophyllaceae menyanthaceae goodeniaceae calyceraceae asteraceae bruniales columelliaceae bruniaceae dipsacales adoxaceae caprifoliaceae escalloniales escalloniaceae paracryphiales paracryphiaceae category taxon identifiers ceratophyllaceae wikidata q12012524 wikispecies ceratophyllaceae apni 54445 col 7wn eol 4343 eppo 1cerf florabase 22812 fna 10178 foao2 ceratophyllaceae foc 10178 gbif old 2425 grin 242 inaturalist 60998 ipni 77126579 1 irmng 119488 itis 18401 nbn nhmsys0021060387 ncbi 32203 nzor 1fb6ca3a 56fa 4ea9 b4c2 d8201239d35f open tree of life 259039 paleobiology database 55409 powo urn lsid ipni org names 77126579 1 tropicos 42000028 vascan 108 vicflora c192d55e 026c 462e b873 d5f040f41b2e watson dallwitz ceratoph wfo wfo 7000000127 worms 394609 retrieved from https en wikipedia org w index php title ceratophyllaceae oldid 1354934229 categories angiosperms freshwater plants angiosperm families hidden categories articles with short description short description is different from wikidata articles with species microformats all articles with dead external links articles with dead external links from june 2024 articles with permanently dead external links taxonbars with 25 29 taxon ids this page was last edited on 19 may 2026 at 01 37 utc page was rendered with parsoid text is available under the creative commons attribution sharealike 4 0 license additional terms may apply by using this site you agree to the terms of use and privacy policy wikipedia is a registered trademark of the wikimedia foundation inc a non profit organization privacy policy about wikipedia disclaimers contact wikipedia legal safety contacts code of conduct developers statistics cookie statement mobile view search search toggle the table of contents ceratophyllaceae 20 languages add topic