Meta tags:

Headings (most frequently used words):

fylax, contents, discovery, and, naming, classification, references,

Text of the page (most frequently used words):
the (43), fylax (20), and (13), from (12), dinosaur (10), clade (10), late (9), cretaceous (9), wikipedia (7), was (7), 2021 (7), edit (7), tethyshadros (6), ornithopoda (6), page (5), taxon (5), hadrosauridae (5), new (5), hadrosauroid (5), prieto (5), márquez (5), carrera (5), farias (5), thyrakolasus (5), dentary (5), genus (5), contents (4), search (4), this (4), non (4), with (4), articles (4), hadrosauroidea (4), telmatosaurus (4), kryptohadros (4), bactrosaurus (4), see (4), below (4), ornithischia (4), doi (4), early (4), 2024 (4), hadrosauromorpha (4), hide (4), move (4), sidebar (4), view (3), using (3), 2026 (3), species (3), description (3), wikidata (3), spain (3), described (3), maastrichtian (3), edmontosaurus (3), hadrosaur (3), lambeosaurinae (3), eotrachodon (3), telmatosauridae (3), tanius (3), protohadros (3), plesiohadros (3), penelopognathus (3), jeyawati (3), gilmoreosaurus (3), eolambia (3), claosaurus (3), dinosauria (3), 2025 (3), formation (3), hadrosaurid (3), diverging (3), longrich (3), matrix (3), sister (3), position (3), classification (3), name (3), keeper (3), tools (3), main (3), languages (2), toggle (2), table (2), contact (2), about (2), privacy (2), policy (2), terms (2), wikimedia (2), commons (2), categories (2), short (2), dinosaurs (2), fossil (2), mummy (2), research (2), contains (2), nomina (2), other (2), related (2), edmontosaurini (2), saurolophini (2), kritosaurini (2), wulagasaurus (2), brachylophosaurini (2), saurolophinae (2), lambeosaurini (2), parasaurolophini (2), tsintaosaurus (2), nipponosaurus (2), jaxartosaurus (2), aralosaurus (2), hadrosaurus (2), zhanghenglong (2), yunganglong (2), shuangmiaosaurus (2), nanyangosaurus (2), nanningosaurus (2), lophorhothon (2), levnesovia (2), huehuecanauhtlus (2), gobihadros (2), comptonatus (2), mantellisaurus (2), barilium (2), emiliasaura (2), kangnasaurus (2), muttaburrasaurus (2), fostoria (2), avemetatarsalia (2), phylum (2), chordata (2), kingdom (2), animalia (2), magyar (2), 2607800 (2), duck (2), billed (2), european (2), archipelago (2), dai (2), hui (2), 105995 (2), bibcode (2), pereda (2), suberbiola (2), scientific (2), albert (2), miguel (2), pdf (2), surviving (2), 1999 (2), references (2), mongoliensis (2), their (2), version (2), initially (2), recovered (2), placed (2), more (2), derived (2), cladogram (2), comes (2), greek (2), hell (2), holotype (2), left (2), figuerola (2), discovery (2), naming (2), appearance (2), upload (2), file (2), changes (2), links (2), history (2), read (2), article (2), log (2), create (2), account (2), donate (2), menu (2), add, topic, mobile, cookie, statement, statistics, developers, code, conduct, legal, safety, contacts, disclaimers, text, available, under, additional, may, apply, site, you, agree, registered, trademark, profit, organization, foundation, inc, use, creative, attribution, sharealike, license, rendered, parsoid, last, edited, march, utc, hidden, microformats, matches, taxa, genera, retrieved, https, org, index, php, title, oldid, 1343737352, 438296, paleobiology, database, old, 11152645, gbif, q107088395, identifiers, r4036, amnh, 5060, timeline, nuda, list, informal, names, hypsilophodont, diet, dakota, basturs, poble, bonebed, arkharavia, willinakaqe, trachodon, tichosteus, thespesius, syngonosaurus, stephanosaurus, siluosaurus, pteropelyx, orthomerus, ornithotarsus, microhadrosaurus, mandschurosaurus, loncosaurus, koutalisaurus, glishades, fulgurotherium, eucercosaurus, diclonius, claorhynchus, cionodon, arstanosaurus, dubia, guegoolithus, ootaxa, wintonopus, tetrapodosaurus, staurichnium, sousaichnium, shiraminesauropus, ornithopodichnites, orcauichnites, iguanodontipus, hadrosaurichnus, dinehichnus, dakotasaurus, caririchnium, camptosaurichnus, bonaparteichnium, anomoepus, amblydactylus, ichnotaxa, shantungosaurus, laiyangosaurus, kundurosaurus, kerberosaurus, kamuysaurus, saurolophus, prosaurolophus, augustynolophus, rhinorex, naashoibitosaurus, kritosaurus, gryposaurus, coahuilasaurus, anasazisaurus, ahshislesaurus, secernosaurus, kelumapusaura, huallasaurus, bonapartesaurus, austrokritosauria, probrachylophosaurus, ornatops, maiasaura, gongshuilong, brachylophosaurus, acristavus, barsboldia, velafrons, olorotitan, magnapaulia, lambeosaurus, hypacrosaurus, corythosaurus, amurosaurus, tlatolophus, plesiolophus, parasaurolophus, charonosaurus, adelolophus, taleta, pararhabdodon, minqaria, latirhinus, kazaklambia, canardia, blasisaurus, arenysaurus, angulomastacator, ajnabia, adynomosaurus, yamatosaurus, malefica, lapampasaurus, aquilarhinus, zuoyunlong, xuwulong, sirindhorna, siamodon, qianjiangsaurus, probactrosaurus, portellsaurus, koshisaurus, jintasaurus, hypsibema, haolong, gonkoken, gongpoquansaurus, equijubus, datonglong, choyrodon, cariocecus, brighstoneus, batyrosaurus, altirhinus, iguanodon, iguanodontidae, ouranosaurus, morelladon, jinzhousaurus, hypselospinus, bolong, hadrosauriformes, uteodon, theiophytalia, riabininohadros, ratchasimasaurus, proa, planicoxa, osmakasaurus, oblitosaurus, napaisaurus, magnamanus, lurdusaurus, lanzhousaurus, istiorachis, iguanacolossus, hippodraco, fukuisaurus, dakotadon, cumnoria, cedrorestes, camptosaurus, calvarius, bayannurosaurus, ankylopollexia, valdosaurus, iyuku, eousdryosaurus, elrhazosaurus, dysalotosaurus, dryosaurus, dryosauridae, owenodon, hesperonyx, draconyx, dryomorpha, zalmoxes, transylvanosaurus, rhabdodon, pareisactus, mochlodon, matheronodon, rhabdodontidae, tenontosaurus, iani, convolosaurus, tenontosauridae, obelignathus, foskeia, ampelognathus, rhabdodontomorpha, weewarrasaurus, trinisaura, tietasaura, talenkauen, sektensaurus, qantassaurus, notohypsilophodon, morrosaurus, mahuidacursor, macrogryphosaurus, leaellynasaura, isasicursor, gasparinisaura, galleonosaurus, diluvicursor, chakisaurus, atlascopcosaurus, anabisetia, elasmaria, vectidromeus, hypsilophodon, gideonmantellia, hypsilophodontidae, thescelosauridae, callovosaurus, burianosaurus, cerapoda, neornithischia, ősi, csiki, sava, budai, botfalvai, 1080, 14772019, journal, systematic, palaeontology, hațeg, basin, densuș, ciula, romania, documents, endemic, hadrosauroids, south, eastern, qingyu, xiong, can, lin, zeng, tan, chao, wang, jun, zhang, yuguang, xing, hai, february, press, 0195, 6671, issn, 1016, cretres, 2025crres, 16605995d, 166, southwest, china, support, for, interchange, faunas, across, east, asia, during, bardet, jalil, 3665, 38351204, pmid, 10864364, pmc, 1038, s41598, 024, 53447, 2024natsr, 3665l, reports, small, duckbilled, morocco, diversity, north, africa, 4202, app, 00821, 2020, acta, palaeontologica, polonica, ibero, armorican, role, biogeography, casanovas, santafé, weishampel, primitive, euhadrosaurian, uppermost, ager, syncline, southern, pyrenees, catalonia, 345, 356, geologie, mijnbouw, corythosauria, arenysaurini, spinorhinus, sachalinensis, aralensis, tuberiferus, dongi, euhadrosauria, foulkii, rugoculus, tiquichensis, yangchengensis, orientalis, weishampeli, djadokhtaensis, insularis, kallaiae, transsylvanicus, agilis, dashiensis, gilmorei, johnsoni, zhugeii, sinensis, transoxiana, datongensis, byrdi, caroljonesa, colleagues, included, updated, similar, but, less, extensive, both, datasets, failed, recover, close, relationship, between, proposed, instead, also, containing, deemed, latter, member, tree, dataset, chinese, these, results, are, displayed, basal, majority, rule, tethyhadros, phylogenetic, making, one, latest, hadrosauromorphs, shown, ángel, belonging, generic, modern, fýlax, specific, thýra, gate, kólasi, thus, creating, combination, gates, reference, proximity, paleogene, mass, extinction, event, ips, 36338, discovered, 1990s, found, northeastern, lleida, province, reconstructed, skeleton, based, preserved, white, meaning, single, known, nearly, complete, ornithopod, binomial, reptilia, class, multiple, angles, temporal, range, preꞓ, 052, period, free, encyclopedia, item, projects, printable, download, print, export, switch, legacy, parser, get, shortened, url, cite, information, permanent, link, what, here, general, actions, english, talk, українська, türkçe, nederlands, français, español, čeština, top, personal, special, pages, recent, community, portal, learn, help, contribute, random, current, events, navigation, jump, content,

Text of the page (random words):
fylax wikipedia jump to content main menu main menu move to sidebar hide navigation main page contents current events random article about wikipedia contact us contribute help learn to edit community portal recent changes upload file special pages search search appearance donate create account log in personal tools donate create account log in contents move to sidebar hide top 1 discovery and naming 2 classification 3 references toggle the table of contents fylax 7 languages čeština español français nederlands türkçe українська 中文 edit links article talk english read edit view history tools tools move to sidebar hide actions read edit view history general what links here related changes upload file permanent link page information cite this page get shortened url switch to legacy parser print export download as pdf printable version in other projects wikimedia commons wikidata item appearance move to sidebar hide from wikipedia the free encyclopedia genus of hadrosauroid dinosaur from the late cretaceous period fylax temporal range late cretaceous 66 052 66 ma preꞓ ꞓ o s d c p t j k pg n holotype dentary from multiple angles scientific classification kingdom animalia phylum chordata class reptilia clade dinosauria clade ornithischia clade ornithopoda clade hadrosauromorpha genus fylax prieto márquez carrera farias 2021 species f thyrakolasus binomial name fylax thyrakolasus prieto márquez carrera farias 2021 fylax meaning keeper is a genus of hadrosauroid ornithopod from the late cretaceous figuerola formation of spain the genus contains a single species fylax thyrakolasus known from a nearly complete left dentary discovery and naming edit reconstructed skeleton based on tethyshadros with the preserved dentary in white the holotype of fylax ips 36338 a left dentary was discovered in the early 1990s it was found in the figuerola formation in lleida province northeastern spain it was initially described in 1999 1 2 in 2021 albert prieto márquez and miguel ángel carrera farias described the dentary as belonging to a new genus of hadrosauroid dinosaur the generic name fylax comes from the modern greek fýlax keeper and the specific name thyrakolasus comes from the greek thýra gate and kólasi hell thus creating the combination keeper of the gates of hell in reference to the proximity of this taxon to the cretaceous paleogene mass extinction event 2 classification edit prieto márquez and carrera farias 2021 recovered fylax as the sister taxon to tethyshadros in a derived position in the hadrosauromorpha making it one of the latest surviving non hadrosaurid hadrosauromorphs their cladogram is shown below hadrosauromorpha jeyawati eolambia protohadros tanius bactrosaurus gilmoreosaurus lophorhothon penelopognathus telmatosaurus claosaurus fylax tethyshadros plesiohadros eotrachodon hadrosauridae in their 2026 description of kryptohadros magyar and colleagues included fylax in an updated version of the phylogenetic matrix of longrich et al 2024 3 and the similar but less extensive matrix of dai et al 2025 4 both datasets failed to recover a close relationship between fylax and tethyshadros as initially proposed in 2021 instead tethyhadros was recovered as the sister taxon of kryptohadros in a clade also containing telmatosaurus deemed the telmatosauridae using the latter matrix fylax was placed in a more derived position as an early diverging member of the hadrosauridae the majority rule tree using the dataset of longrich et al 2024 placed fylax in a more basal position as the sister taxon to the chinese bactrosaurus these results are displayed in the cladogram below 5 hadrosauromorpha eolambia caroljonesa protohadros byrdi yunganglong datongensis levnesovia transoxiana tanius sinensis nanyangosaurus zhugeii bactrosaurus johnsoni fylax thyrakolasus gilmoreosaurus mongoliensis shuangmiaosaurus gilmorei nanningosaurus dashiensis claosaurus agilis telmatosauridae telmatosaurus transsylvanicus kryptohadros kallaiae tethyshadros insularis plesiohadros djadokhtaensis gobihadros mongoliensis penelopognathus weishampeli eotrachodon orientalis zhanghenglong yangchengensis huehuecanauhtlus tiquichensis jeyawati rugoculus hadrosauridae hadrosaurus foulkii euhadrosauria saurolophinae wulagasaurus dongi brachylophosaurini kritosaurini edmontosaurini saurolophini lambeosaurinae aralosaurus tuberiferus jaxartosaurus aralensis nipponosaurus sachalinensis tsintaosaurus spinorhinus arenysaurini corythosauria lambeosaurini parasaurolophini references edit casanovas m l pereda suberbiola x p santafé j v and weishampel d b 1999 a primitive euhadrosaurian dinosaur from the uppermost cretaceous of the ager syncline southern pyrenees catalonia geologie en mijnbouw 78 345 356 1 2 prieto márquez albert carrera farias miguel 2021 a new late surviving early diverging ibero armorican duck billed dinosaur and the role of the late cretaceous european archipelago in hadrosauroid biogeography pdf acta palaeontologica polonica 66 doi 10 4202 app 00821 2020 longrich n r pereda suberbiola x bardet n jalil n e 2024 a new small duckbilled dinosaur hadrosauridae lambeosaurinae from morocco and dinosaur diversity in the late maastrichtian of north africa scientific reports 14 1 3665 bibcode 2024natsr 14 3665l doi 10 1038 s41598 024 53447 9 pmc 10864364 pmid 38351204 dai hui ma qingyu xiong can lin yu zeng hui tan chao wang jun zhang yuguang xing hai february 2025 a new late diverging non hadrosaurid hadrosauroid dinosauria ornithopoda from southwest china support for interchange of dinosaur faunas across east asia during the late cretaceous cretaceous research 166 in press 105995 bibcode 2025crres 16605995d doi 10 1016 j cretres 2024 105995 issn 0195 6671 magyar j ősi a csiki sava z budai s botfalvai g 2026 new early maastrichtian duck billed dinosaur from hațeg basin densuș ciula formation romania documents an endemic clade of non hadrosaurid hadrosauroids in the south eastern late cretaceous european archipelago journal of systematic palaeontology 24 1 2607800 doi 10 1080 14772019 2025 2607800 v t e ornithopoda kingdom animalia phylum chordata clade dinosauria clade ornithischia clade neornithischia clade cerapoda avemetatarsalia see avemetatarsalia ornithischia see ornithischia ornithopoda see below ornithopoda ornithopoda burianosaurus callovosaurus thescelosauridae hypsilophodontidae gideonmantellia hypsilophodon vectidromeus elasmaria anabisetia atlascopcosaurus chakisaurus diluvicursor fostoria galleonosaurus gasparinisaura isasicursor kangnasaurus leaellynasaura macrogryphosaurus mahuidacursor morrosaurus muttaburrasaurus notohypsilophodon qantassaurus sektensaurus talenkauen tietasaura trinisaura weewarrasaurus rhabdodontomorpha ampelognathus emiliasaura foskeia fostoria muttaburrasaurus obelignathus tenontosauridae convolosaurus iani tenontosaurus rhabdodontidae matheronodon mochlodon pareisactus rhabdodon transylvanosaurus zalmoxes dryomorpha draconyx hesperonyx owenodon dryosauridae dryosaurus dysalotosaurus elrhazosaurus eousdryosaurus iyuku kangnasaurus valdosaurus ankylopollexia barilium bayannurosaurus calvarius camptosaurus cedrorestes cumnoria dakotadon emiliasaura fukuisaurus hippodraco iguanacolossus istiorachis lanzhousaurus lurdusaurus magnamanus napaisaurus oblitosaurus osmakasaurus planicoxa proa ratchasimasaurus riabininohadros theiophytalia uteodon hadrosauriformes bolong hypselospinus jinzhousaurus mantellisaurus morelladon ouranosaurus iguanodontidae barilium comptonatus iguanodon mantellisaurus hadrosauroidea see below hadrosauroidea hadrosauroidea altirhinus bactrosaurus batyrosaurus brighstoneus cariocecus choyrodon claosaurus comptonatus datonglong eolambia equijubus fylax gilmoreosaurus gobihadros gongpoquansaurus gonkoken haolong huehuecanauhtlus hypsibema jeyawati jintasaurus koshisaurus levnesovia lophorhothon nanningosaurus nanyangosaurus penelopognathus plesiohadros portellsaurus probactrosaurus protohadros qianjiangsaurus shuangmiaosaurus siamodon sirindhorna tanius xuwulong yunganglong zhanghenglong zuoyunlong telmatosauridae kryptohadros telmatosaurus tethyshadros hadrosauridae aquilarhinus eotrachodon hadrosaurus lapampasaurus malefica yamatosaurus lambeosaurinae adynomosaurus ajnabia angulomastacator aralosaurus arenysaurus blasisaurus canardia jaxartosaurus kazaklambia latirhinus minqaria nipponosaurus pararhabdodon taleta tsintaosaurus parasaurolophini adelolophus charonosaurus parasaurolophus plesiolophus tlatolophus lambeosaurini amurosaurus corythosaurus hypacrosaurus lambeosaurus magnapaulia olorotitan velafrons saurolophinae barsboldia brachylophosaurini acristavus brachylophosaurus gongshuilong maiasaura ornatops probrachylophosaurus wulagasaurus austrokritosauria bonapartesaurus huallasaurus kelumapusaura secernosaurus kritosaurini ahshislesaurus anasazisaurus coahuilasaurus gryposaurus kritosaurus naashoibitosaurus rhinorex saurolophini augustynolophus prosaurolophus saurolophus edmontosaurini edmontosaurus kamuysaurus kerberosaurus kundurosaurus laiyangosaurus shantungosaurus related articles ichnotaxa amblydactylus anomoepus bonaparteichnium camptosaurichnus caririchnium dakotasaurus dinehichnus hadrosaurichnus iguanodontipus orcauichnites ornithopodichnites shiraminesauropus sousaichnium staurichnium tetrapodosaurus wintonopus ootaxa guegoolithus nomina dubia arstanosaurus cionodon claorhynchus diclonius eucercosaurus fulgurotherium glishades koutalisaurus loncosaurus mandschurosaurus microhadrosaurus ornithotarsus orthomerus pteropelyx siluosaurus stephanosaurus syngonosaurus thespesius tichosteus trachodon willinakaqe other articles arkharavia basturs poble bonebed dakota fossil hadrosaur diet hypsilophodont list of informal dinosaur names contains hadrosaur nomina nuda timeline of hadrosaur research edmontosaurus mummy amnh 5060 edmontosaurus mummy s m r4036 taxon identifiers fylax wikidata q107088395 gbif old 11152645 paleobiology database 438296 retrieved from https en wikipedia org w index php title fylax oldid 1343737352 categories hadrosauroidea dinosaur genera maastrichtian dinosaurs fossil taxa described in 2021 dinosaurs of spain hidden categories articles with short description short description matches wikidata articles with species microformats this page was last edited on 16 march 2026 at 02 24 utc page was rendered with parsoid text is available under the creative commons attribution sharealike 4 0 license additional terms may apply by using this site you agree to the terms of use and privacy policy wikipedia is a registered trademark of the wikimedia foundation inc a non profit organization privacy policy about wikipedia disclaimers contact wikipedia legal safety contacts code of conduct developers statistics cookie statement mobile view search search toggle the table of contents fylax 7 languages add topic