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al parts conceded that it would be rash to rule out the possibility that trade in synthetic rhinoceros horn could play a role in future conservation strategies 143 historical representations upper paleolithic cave paintings of woolly rhinoceros at chauvet cave france replica woolly rhinoceroses are depicted the european paleolithic art such as in cave paintings in chauvet cave in france 144 which date to around 30 40 000 years ago 145 western zhou bronze rhino greek historian and geographer agatharchides 2nd century bc mentions the rhinoceros in his book on the erythraean sea 146 in khmer art the hindu god agni is depicted with a rhinoceros as his vahana 147 148 similarly in medieval era thai literature agni also called phra phloeng is sometimes described as riding a rhinoceros 149 150 albrecht dürer created a famous woodcut of a rhinoceros in 1515 based on a written description and brief sketch by valentim fernandes a german printer resident in lisbon 151 he never saw the animal itself so dürer s rhinoceros is a somewhat inaccurate depiction 152 there are legends about rhinoceroses stamping out fire in burma india and malaysia the mythical rhinoceros has a special name in malay badak api wherein badak means rhinoceros and api means fire the animal would come when a fire was lit in the forest and stamp it out 153 there are no recent confirmations of this phenomenon this legend was depicted in the film the gods must be crazy 1980 which shows an african rhinoceros putting out two campfires 154 in 1974 a lavender rhinoceros symbol began to be used as a symbol of the gay community in boston united states 155 rhinoceroses in art a wine vessel in the form of a bronze rhinoceros with silver inlay from the western han 202 bc 9 ad period of china sporting a saddle on its back dürer s rhinoceros an albrecht dürer woodcut from 1515 a rhinoceros depicted on a roman mosaic in villa romana del casale an archeological site near piazza armerina in sicily italy how the rhinoceros got his skin from rudyard kipling s just so stories from 1902 a lavender rhinoceros a symbol used as a sign of gay visibility see also conservation bardiya national park chitwan national park care for wild rhino sanctuary international rhino foundation kaziranga national park list of odd toed ungulates by population nicolaas jan van strien save the rhino traffic individual rhinoceroses abada clara list of fictional ungulates rhinoceros of versailles literature rhinoceros 1959 play other rhinoceroses in ancient china references glossary american museum of natural history archived from the original on 20 november 2021 osborn henry fairfield 1898 the extinct rhinoceroses memoirs of the american museum of natural history 1 3 80 sanisidro oscar cantalapiedra juan l 2022 the rhinocerotidae fossil record in the iberian peninsula historical biology 34 8 1591 1610 doi 10 1080 08912963 2022 2051502 issn 0891 2963 owen smith norman 1984 macdonald d ed the encyclopedia of mammals new york facts on file pp 490 495 isbn 978 0 87196 871 5 vietnam s appetite for rhino horn drives poaching in africa npr retrieved 27 september 2021 what is a rhinoceros horn made of yesmag bc ca 9 october 2003 archived from the original on 28 september 2011 retrieved 23 september 2010 cheung hubert mazerolle lorraine possingham hugh biggs duan 1 february 2021 rhino horn use by consumers of traditional chinese medicine in china conservation science and practice 3 5 e365 bibcode 2021consp 3e 365c doi 10 1111 csp2 365 hdl 11573 1713877 1 2 vigne lucy martin esmond january march 2018 amid conflict yemen s demand for rhino horn daggers continues pdf swara archived pdf from the original on 7 march 2023 via rhino resource center tougard c et al 2001 phylogenetic relationships of the five extant rhinoceros species rhinocerotidae perissodactyla based on mitochondrial cytochrome b and 12s rrna genes definition of rhinoceros merriam webster dictionary 28 august 2023 retrieved 23 september 2023 rhinoceros dictionary cambridge org 26 november 2025 retrieved 1 december 2025 rookmaaker l c april 2005 review of the european perception of the african rhinoceros journal of zoology 265 4 365 376 doi 10 1017 s0952836905006436 issn 0952 8369 rookmaaker kees 2015 rhino systematics in the times of linnaeus cuvier gray and groves in behie alison m oxenham marc f eds taxonomic tapestries the threads of evolutionary behavioural and conservation research anu press pp 299 320 isbn 978 1 925022 36 0 jstor j ctt169wd9c 18 retrieved 1 december 2025 1 2 giaourtsakis ioannis x 2022 the fossil record of rhinocerotids mammalia perissodactyla rhinocerotidae in greece in vlachos evangelos ed fossil vertebrates of greece vol 2 cham springer international publishing pp 409 500 doi 10 1007 978 3 030 68442 6_14 isbn 978 3 030 68441 9 s2cid 239883886 retrieved 20 november 2023 gray je 1821 on the natural arrangement of vertebrose animals london medical repository 15 297 310 rhinocerotidae mammal species of the world 3rd ed retrieved 2 december 2025 owen r 1845 odontography london uk hippolyte bailliere vol 1 p 587 tissier jérémy antoine pierre olivier becker damien july 2020 new material of epiaceratherium and a new species of mesaceratherium clear up the phylogeny of early rhinocerotidae perissodactyla royal society open science 7 7 200633 bibcode 2020rsos 700633t doi 10 1098 rsos 200633 issn 2054 5703 pmc 7428265 pmid 32874655 1 2 3 mallet christophe houssaye alexandra cornette raphaël billet guillaume 2 november 2022 long bone shape variation in the forelimb of rhinocerotoidea relation with size body mass and body proportions zoological journal of the linnean society 196 3 1201 1234 doi 10 1093 zoolinnean zlab095 issn 0024 4082 mihlbachler matthew c 15 november 2007 sexual dimorphism and mortality bias in a small miocene north american rhino menoceras arikarense insights into the coevolution of sexual dimorphism and sociality in rhinos journal of mammalian evolution 14 4 217 238 doi 10 1007 s10914 007 9048 4 issn 1064 7554 1 2 3 4 d r prothero rhinocerotidae c m janis k m scott l jacobs eds evolution of tertiary mammals of north america cambridge university press cambridge 1998 pp 595 605 hillman smith a k k owen smith n anderson j l hall martin a j selaladi j p november 1986 age estimation of the white rhinoceros ceratotherium simum journal of zoology 210 3 355 377 doi 10 1111 j 1469 7998 1986 tb03639 x issn 0952 8369 boy s c raubenheimer e j marais j steenkamp g july 2015 white rhinoceros ceratotherium simum horn development and structure a deceptive optical illusion journal of zoology 296 3 161 166 doi 10 1111 jzo 12229 issn 0952 8369 1 2 3 4 lu xiao kang deng tao pandolfi luca 16 february 2023 reconstructing the phylogeny of the hornless rhinoceros aceratheriinae frontiers in ecology and evolution 11 1005126 bibcode 2023freev 1105126l doi 10 3389 fevo 2023 1005126 issn 2296 701x hieronymus t l 2009 biological sciences osteological correlates of cephalic skin structures in amniota documenting the evolution of display and feeding structures with fossil data phd thesis athens college of arts and sciences of ohio university usa p 33 43 van dongen pam 1998 brain size in vertebrates in the central nervous system of vertebrates eds nieuwenhuys r ten donkelaar hj nicholson c pp 2099 2134 berlin germany springer 1 2 bhagwandin adhil haagensen mark manger paul r 31 august 2017 the brain of the black diceros bicornis and white ceratotherium simum african rhinoceroses morphology and volumetrics from magnetic resonance imaging frontiers in neuroanatomy 11 74 doi 10 3389 fnana 2017 00074 issn 1662 5129 pmc 5583206 pmid 28912691 shadwick r e russell a p lauff r f 29 september 1992 the structure and mechanical design of rhinoceros dermal armour philosophical transactions of the royal society of london series b biological sciences 337 1282 419 428 doi 10 1098 rstb 1992 0118 issn 0962 8436 pmid 1359589 endo hideki kobayashi hiroshi koyabu daisuke hayashida akiko jogahara takamichi taru hajime oishi motoharu itou takuya koie hiroshi sakai takeo december 2009 the morphological basis of the armor like folded skin of the greater indian rhinoceros as a thermoregulator mammal study 34 4 195 200 doi 10 3106 041 034 0403 issn 1343 4152 groves colin p leslie david m 21 january 2011 rhinoceros sondaicus perissodactyla rhinocerotidae mammalian species 43 190 208 bibcode 2011mamsp 43 190g doi 10 1644 887 1 hdl 1885 52865 issn 0076 3519 succop nick dicerorhinus sumatrensis sumatran rhinoceros animal diversity web retrieved 26 december 2024 houck ml ryder oa váhala j kock ra oosterhuis je january february 1994 diploid chromosome number and chromosomal variation in the white rhinoceros ceratotherium simum the journal of heredity 85 1 30 34 pmid 8120356 1 2 3 hutchins m kreger m d july 2006 rhinoceros behaviour implications for captive management and conservation international zoo yearbook 40 1 150 173 doi 10 1111 j 1748 1090 2006 00150 x issn 0074 9664 luo yu heng wright andré denis g li you long li hua yang qi hong luo ling juan yang ming xian 2013 diversity of methanogens in the hindgut of captive white rhinoceroses ceratotherium simum bmc microbiology 13 1 207 doi 10 1186 1471 2180 13 207 issn 1471 2180 pmc 3846858 pmid 24228793 schwarzenberger franz hermes robert april 2023 comparative analysis of gestation in three rhinoceros species diceros bicornis ceratotherium simum rhinoceros unicornis general and comparative endocrinology 334 114214 doi 10 1016 j ygcen 2023 114214 pmid 36646327 1 2 muntifering jeff r guerier abigail beytell piet stratford ken september 2023 population parameters performance and insights into factors influencing the reproduction of the black rhinoceros diceros bicornis in namibia oryx 57 5 659 669 doi 10 1017 s0030605322001065 issn 0030 6053 roth terri l reinhart paul r romo john s candra dedi suhaery andriansyah stoops monica a september 2013 sexual maturation in the sumatran rhinoceros dicerorhinus sumatrensis sumatran rhino sexual maturation zoo biology 32 5 549 555 doi 10 1002 zoo 21089 pmid 23897764 cinková ivana policht richard january 2015 discrimination of familiarity and sex from chemical cues in the dung by wild southern white rhinoceros animal cognition 18 1 385 392 doi 10 1007 s10071 014 0810 8 issn 1435 9448 pmid 25273823 1 2 skinner john d chimimba christian t 2005 the mammals of the southern african subregion cambridge university press p 527 isbn 978 0 521 84418 5 rookmaaker kees 2003 why the name of the white rhinoceros is not appropriate pachyderm 34 88 93 doi 10 69649 pachyderm v34i1 1137 archived from the original on 18 august 2022 retrieved 24 february 2012 the white rhinoceros bisbee s fish wildlife conservation fund retrieved 24 november 2024 robinson terry j v trifonov i espie e h harley january 2005 interspecific hybridization in rhinoceroses confirmation of a black white rhinoceros hybrid by karyotype fluorescence in situ hybridization fish and 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raymond williams amirtharaj christy nathan senthival ramono widodo s clements gopalasamy reuben 2011 now or never what will it take to save the sumatran rhinoceros dicerorhinus sumatrensis from extinction oryx 45 2 225 233 doi 10 1017 s0030605310000864 issn 0030 6053 benoît goossens milena salgado lynn jeffrine j rovie ryan abdul h ahmad junaidi payne zainal z zainuddin senthilvel k s s nathan laurentius n ambu 2013 genetics and the last stand of the sumatran rhinoceros dicerorhinus sumatrensis oryx 47 3 340 344 doi 10 1017 s0030605313000045 beachy ben 7 december 2015 sharks tigers and elephants new analysis reveals tpp threats to endangered species sierra club wang haibing bai bin meng jin wang yuanqing 21 december 2016 earliest known unequivocal rhinocerotoid sheds new light on the origin of giant rhinos and phylogeny of early rhinocerotoids scientific reports 6 1 39607 bibcode 2016natsr 639607w doi 10 1038 srep39607 issn 2045 2322 pmc 5175171 pmid 28000789 antoine p o ducrocq s marivaux l chaimanee y crochet j y jaeger j j welcomme j l 2003 early rhinocerotids mammalia perissodactyla from south asia and a review of the holarctic paleogene rhinocerotid record can j earth sci 40 365 374 1 2 tissier jérémy antoine pierre olivier becker damien july 2020 new material of epiaceratherium and a new species of mesaceratherium clear up the phylogeny of early rhinocerotidae perissodactyla royal society open science 7 7 200633 bibcode 2020rsos 700633t doi 10 1098 rsos 200633 issn 2054 5703 pmc 7428265 pmid 32874655 1 2 cerdeño esperanza august 1998 diversity and evolutionary trends of the family rhinocerotidae perissodactyla palaeogeography palaeoclimatology palaeoecology 141 1 2 13 34 bibcode 1998ppp 141 13c doi 10 1016 s0031 0182 98 00003 0 1 2 3 4 liu shanlin westbury michael v dussex nicolas mitchell kieren j sinding mikkel holger s heintzman peter d duchêne david a kapp joshua d von seth johanna heiniger holly sánchez barreiro fátima 24 august 2021 ancient and modern genomes unravel the evolutionary history of the r...
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